1. Phonekeo, S., Mlot, N., Monaenkova, D., Hu, D.L., Tovey, C. (2017) Fire ants perpetually rebuild sinking towers. Soc. open sci. 2017 4 170475; DOI: 10.1098/rsos.170475.   pdf
  2. Noel, A., Guo, H.-Y., Mandica, M., Hu, D.L. (2017) Frogs use a viscoelastic tongue and non-Newtonian saliva to catch prey. Journal of the Royal Society Interface. 2017 14 20160764; DOI: 10.1098/rsif.2016.0764. pdf
  3. Amador, Guillermo J., and David L. Hu. “Sticky Solution Provides Grip for the First Robotic Pollinator.” Chem 2.2 (2017): 162-164. pdf
  4. Amador, G., Matherne, M., Waller, D., Mathews, M., Gorb, S., Hu, D.L. (2017). Honeybee hairs are essential for pollen capture and removal. Bioinspiration & Biomimetics12 (2): 026015. https://doi.org/10.1088/1748-3190/aa5c6e pdf
  5. Yang, P.J., LaMarca, M., Kaminski, C., Chu, D., Hu, D.L. (2017) The Hydrodynamics of Defecation. Soft Matter.  pdf


  1. Tennebaum, M., Liu, Z., Hu, D.L. Fernandez-Nieves, A. (2016) Mechanics of fire ant aggregations.  Nature Materials. 15, 54–59 doi:10.1038/nmat4450 pdf
  2. Hu, D. L., Lefton, L., & Ludovice, P. J. (2016). Humour Applied to STEM Education. Systems Research and Behavioral Sciencepdf
  3. Marvi, H., Cook, J. P., Streator, J. L., & Hu, D. L. (2016). Snakes move their scales to increase friction. Biotribology, 5, 52-60. pdf
  4. Hu, D. L., Phonekeo, S., Altshuler, E., & Brochard-Wyart, F. (2016). Entangled active matter: From cells to ants. The European Physical Journal Special Topics, 225(4), 629-649. pdf
  5. Phonekeo, S., Dave, T., Kern, M., Franklin, S.V. and Hu, D.L. (2016). Ant aggregations self-heal to compensate for Ringlemann Effect. Soft Matter. 12(18) :4214-20. doi: 10.1039/c6sm00063k.pdf
  6. Hu, D.L. (2016) A Fish Grew Legs and Other Tales. BioScience 2016; 66 (9): 792-793. doi: 10.1093/biosci/biw094
  7. Hu, D.L. (2016) Confessions of a Wasteful Scientist. Scientific American link


  1. Dickerson, A.K, Shankles, P.G., Berry Jr, B.E., Hu, D.L. (2015) Fog and dense gas disrupt mosquito flight due to increased aerodynamic drag on halters. Journal of Fluids and Structures.   10.1016/j.jfluidstructs.2015.03.016 pdf
  2. Amador, G., Mao, W., DeMercurio, P., Montero, C., Clewis, J., Alexeev, A., Hu, D.L. (2015) Eyelashes divert airflow to protect the eye.   Journal of the Royal Society Interface. 20141294. pdf
  3.  Dickerson, A.K., Liu, X., Zhu, T., Hu, D.L. Fog spontaneously folds mosquito wings. (2015) Physics of Fluids 27, 021901 pdf
  4. Guillermo, A.J., Durand, F., Mao, W., Sindhu, P, Hidetoshi, T., Vinh-Thanh, N., Shimayama, I., Alexeev, A., Hu, D.L. (2015) Insects have hairy eyes that reduce particle deposition.  European Physical Journal: Special Topics. 224, 3361–3377 pdf
Guillermo, A.J. and Hu, D.L. (2015) Cleanliness is next to godliness: mechanisms for staying clean.  Journal of Experimental Biology. pdf


  1. Yang, P.J., Pham, J., Choo, J., Hu, D.L. (2014) Duration of urination does not change with body size. Proceedings of the National Academy of Sciences, USA. pdf
  2. Dickerson, A.D., Hu, D.L. (2014) Mosquitoes actively remove drops deposited by fog and dew. Integrative and Comparative Biology. doi: 10.1093/icb/icu042 pdf
  3. Foster, P. C., Mlot, N. J., Lin, A. and Hu, D. L. (2014). Fire ants actively control spacing and orientation within self-assemblages. Journal of Experimental Biology 217, 2089-2100. pdf
  4. Dickerson, A.D., Shankles, P., Hu, D. L. (2014) “Raindrops push and splash flying insects.” Physics of Fluids. 26, 027104. pdf
  5. Levy, R., Lichtman, F., Hu, D.L. (2014) The Scientist-Reporter Collaboration: A Scientist’s Guide to Working with the Press. Society of Industrial and Applied Mathematics (SIAM) News, Volume 47, Number 3, April 2014, pages 5-7. pdf


  1. • Marvi, H, Bridges, J., Hu, D.L. (2013) “Snakes mimic earthworms: Propulsion using Rectilinear Traveling Waves, ” Journal of the Royal Society Interface. 10: 20130188 pdf
  2. • Amador, G., Yamada, Y., McCurley, M., Hu, D. L. (2013) Raindrop acceleration in splash-cup seed dispersal.  Proceedings of the Royal Society: Interface. 10: 20120880. pdf


  1. Dickerson, A., Mills, Z. & Hu, D. L. (2012) Wet mammals shake at tuned frequencies to dry. Journal of the Royal Society Interface pdf
  2.  Dickerson, A., Shankles, P., Madhaven, N., Hu, D. L. (2012) Mosquitoes survive raindrop collisions by virtue of their low mass. Proceedings of the National Academy of Sciences, USA. 109 (25) 9822-9827. pdf
  3.  Marvi, H. & Hu, D. L. (2012) Friction enhancement in concertina locomotion of snakes.  Journal of the Royal Society Interface. pdf
  4.  Gravish, N., Franklin, S.V., Hu, D.L. & Daniel I. Goldman. (2012) Entangled granular media.  Physical Review Letters. 108, 208001 (journal cover). pdf
  5.  West, D., Lal, I., Leamy, M. & Hu, D.L. (2012) Locomotion of Mexican Jumping Beans. Bioinspiration and Biomimetics.  7: 036014 (12pp).  pdf
  6.  Hu, D.L. (2012) Electromechanics for the Twenty-first century.  Bioscience.  62 (1): 94-95.  pdf
  7.  Hobbs, W. & Hu, D.L. (2012) Tree-inspired Piezoelectric Energy Harvesting.  Journal of Fluids and Structures. 20: 103-114. pdf


  1. Marvi, H., Meyers, G., Russell, G. & Hu, D.L. (2011) Scalybot: a snake-inspired robot with active frictional anisotropy. ASME Dynamic Systems and Control Conference Proceedings. pdf
  2. Niu, J & Hu, D.L. (2011) Drag reduction of a hairy disk. Physics of Fluids.  23: 101701 (4 pages). pdf
  3.  Hu, D.L., Sielert, K., Gordon, M. (2011) The strength of turtle shells and mammal skulls. Journal of the Mechanics of Materials and Structures. 6 (9-10): 1197-1211. pdf
  4. Mlot, N.J., Tovey, C.A, Hu, D. L. (2011).  Proceedings of the National Academy of Sciences, USA. 108: 7669-7673.  pdf
  5.  Hu, D. L. Richards, P. & Alexeev, A. (2011).  Mechanics of giant pumpkins: how extreme weight determines shape.  International Journal of Nonlinear Science.  pdf

2009 – 2010

  1. Goldman, D. I. & Hu, D. L. (2010). Wiggling through the world. American Scientist 98, 314–323.  pdf
  2. Bush, J. W. M. & Hu, D. L. (2010). Walking on water. Physics Today 63, 62–63.  pdf • Hu, D. L. & Bush, J. W. M. (2010). The hydrodynamics of water-walking arthropods. Journal of Fluid Mechanics 644: 5-33.  pdf
  3.  Hu, D.L., Nirody, J., Scott, T., Shelley, M.J. (2009). The mechanics of slithering locomotion.  Proceedings of the National Academy of Sciences, USA. 106, 10081-10085.  pdf

2006 – 2008

  1. Bush, J.W.M., Hu, D.L. and Prakash, M. (2008). The Integument of Water-walking Arthropods: Form and Function, Advances in Insect Physiology, 34, 117-192.  pdf
  2. Hu, D.L., Goreau, T., Bush, J.W.M. (2008). Flow visualization using tobacco mosaic virus. Experiments in Fluids. 46: 477-484.  pdf
  3. Hu, D.L., Prakash, M., Chan, B. and Bush, J.W.M. (2007). Water-walking devices, Experiments in Fluids. 43:769–778.   pdf
  4. Bush J.W.M., Hu D.L. (2006). Walking on water: Biolocomotion at the interface.  Annual Review of Fluid Mechanics. 38: 339-369.   pdf

2001 – 2005 

  1. Hu, DL., Bush J.W.M. (2005). Meniscus-climbing insects. Nature. 437: 733-736. pdf
  2. Hu, D.L., Mendel L., Chan B., Goreau T., Bush, J.W.M. (2005). Visualization of a fish wake using tobacco mosaic virus. Physics of Fluids. 17: 091103.   pdf
  3. Hu, D.L., Chan B., Bush J.W.M. (2003). The hydrodynamics of water strider locomotion. Nature. 424: 663-666.  pdf
  4. Hu, D.L., Chan B, Bush J.W.M.  (2003). Water-walking. Physics of Fluids. 15 (9): S10.  pdf